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- January 27, 2020.
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- 1MRC Centre for Global Infectious Disease Analysis, Imperial College London, London, W2 1PG, UK
- 2Department of Zoology, University of Oxford, Oxford, OX1 3SY, UK
- 3Mathematical Sciences, University of Southampton, Highfield, Southampton SO17 1BJ, UK
- ↵* Corresponding author; email: k.parag{at}imperial.ac.uk