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- April 6, 2017.
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- 1Department of Biosystems Science and Engineering, Eidgenössische Tech-nische Hochschule Zurich, 4058 Basel, Switzerland
- 2Swiss Institute of Bioinformatics (SIB), Switzerland
- 3Division of Ecology and Evolution, Research School of Biology, Australian National University, Canberra, Australia
- 4Centre for Computational Evolution, University of Auckland, Auckland, New Zealand
- 5Department of Computer Science, University of Auckland, Auckland, New Zealand
- ↵*Corresponding author: E-mail: tanja.stadler{at}bsse.ethz.ch or chi.zhang{at}bsse.ethz.ch