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- October 17, 2016.
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- 1Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, United Kingdom
- 2Department of Biology, Center for Applied Tropical Ecology and Conservation, University of Puerto Rico, Rio Piedras, Puerto Rico.
- 3Smithsonian Tropical Research Institute, Apartado 0843-03092, Panamá, Panama.
- ↵*Corresponding Author: shm45{at}cam.ac.uk