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- August 10, 2016.
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- 1Wolfson Centre for Mathematical Biology, Mathematical Institute, University of Oxford, Radcliffe Observatory Quarter, Woodstock Road, Oxford, OX2 6GG
- 2MRC Human Genetics Unit, MRC IGMM, Western General Hospital, University of Edinburgh, Edinburgh, EH4 2XU
- 3Centre for Mathematical Biology, Department of Mathematical Sciences, University of Bath, Claverton Down, Bath, BA2 7AY