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- July 28, 2016.
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- 1Milner Centre for Evolution, University of Bath.
- 2University College London. Dept. Genetics, Evolution and Environment. London, UK.
- 3Centre for Mathematics and Physics in the Life Sciences and Experimental Biology (CoMPLEX), University College London. London, UK.
- 4University of Bristol, School of Social and Community Medicine, Bristol, UK.
- ↵*To whom correspondence should be addressed, Milner Centre for Evolution, Department of Biology & Biotechnology, University of Bath, 4 South, Lab 0.39, Claverton Down, Bath, BA2 7AY, United Kingdom, danielfalush{at}googlemail.com