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- September 7, 2015.
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- 1Centre for Mathematics and Physics in the Life Sciences and Experimental Biology, University College London, Gower Street, London WC1E 6BT, United Kingdom
- 2Research Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, United Kingdom
- ↵* Correspondence to: ucbprad{at}ucl.ac.uk