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- May 31, 2015.
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- 1Department of Evolution and Ecology, University of California, Davis, Davis, CA, U.S.A.
- 2Department of Integrative Biology, University of California, Berkeley, CA, 94720, U.S.A.
- 3Department of Statistics, University of California, Berkeley, CA, 94720, U.S.A.
- 4Department of Mathematics, Stockholm University, Stockholm, SE-106 91, Sweden
- Corresponding author: Sebastian Höhna, Department of Integrative Biology, University of California, Berkeley, CA, 94720, U.S.A.; E-mail: Sebastian.Hoehna{at}gmail.com.
↵☯ Equal priority authorship