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- April 15, 2015.
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- 1Dept. Genetics, Evolution & Environment, University College London, Gower Street, London, UK, WC1E 6BT
- 2Dept. Zoology, University of Cambridge, Downing Street, Cambridge, UK, CB2 3EJ
- 3Smithsonian Tropical Research Institute, MRC 0580-12, Unit 9100 Box 0948, DPO AA 34002-9998, Panama
- 4Dept. Biology, University of Leicester, Adrian Building, University Road, Leicester, UK, LE1 7RH
- Corresponding author: Stephen H. Montgomery: stephen.montgomery{at}cantab.net