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- December 4, 2014.
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- 1Department of Evolutionary Theory, Max Planck Institute for Evolutionary Biology, August-Thienemann-Str. 2, 24306 Plön, Germany e-mail: pena{at}evolbio.mpg.de
- 2Faculty of Business and Economics, University of Basel, Peter Merian-Weg 6, CH-4002 Basel, Switzerland e-mail: georg.noeldeke{at}unibas.ch
- 3Department of Ecology and Evolution, University of Lausanne, Le Biophore, CH-1015 Lausanne, Switzerland e-mail: laurent.lehmann{at}unil.ch
- ↵*Corresponding author.