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- October 14, 2014.
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- *Department of Biology, Indiana University, Bloomington, IN 47405, USA
- §School of Informatics and Computing, Indiana University, Bloomington, IN 47405, USA
- 1Current address: Department of Evolution and Ecology, University of California, Davis, CA 95616, USA
- 2Current address: School of Biological Sciences, University of Nebraska, Lincoln, NE 68588
- Author for correspondence: Kristi Montooth, School of Biological Sciences, 1104 T Street, Lincoln, NE 68588-0118, kmontooth2{at}unl.edu