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- September 13, 2014.
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- 1Department of Biology, Stanford University, Stanford, CA 94305-5020
- 2Institut für Populationsgenetik, Vetmeduni Vienna, Veterinärplatz 1, Vienna A-1210, Austria
- 3Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra). Passeig Maritim de la Barceloneta 37-49. 08003 Barcelona, Spain
- 4Department of Biology, The University of Pennsylvania, Philadelphia, PA 19104
- Contact information for corresponding author: Alan O. Bergland Department of Biology Stanford University Stanford, CA 94305-5020 email: bergland{at}stanford.edu phone: (650) 736 2249 fax: (650) 723 6132